Sawa et al. (2007); CO; 16L:8D; WT, gi

Property Value
BioDare ID 13790763084422
Author Sawa, M.
Institution Scripps Research Institute
License CC_BY

Description

Purpose

Precise timing of CONSTANS (CO) gene expression is necessary for day-length discrimination for photoperiodic flowering. The FLAVIN-BINDING, KELCH REPEAT, F-BOX 1 (FKF1), and GIGANTEA (GI) proteins regulate CO transcription in Arabidopsis. We demonstrate that FKF1 and GI proteins form a complex in a blue-light–dependentmanner. The timing of this interaction regulates the timing of daytime CO expression. FKF1 function is dependent on GI, which interacts with a CO repressor, CYCLING DOF FACTOR 1 (CDF1), and controls CDF1 stability. GI, FKF1, and CDF1 proteins associate with CO chromatin. Thus, the FKF1-GI complex forms on the CO promoter in late afternoon to regulate CO expression, providing a mechanistic view of how the coincidence of light with circadian timing regulates photoperiodic flowering.

Comments

Sample preparation

To quantify mRNA of genes involved in flowering regulation, total RNA was isolated from 10-day-old seedlings using RNeasy plant mini kit (Qiagen). For synthesis of cDNA, 2.5 μg of total RNA was reverse-transcribed with the iScript cDNA synthesis kit (Bio-Rad). The synthesized cDNAs were amplified by using a Bio-Rad real-time detection system. The primers used for amplification of CO, FKF1 and GI were described previously (Nakamichi et al., 2007). The forward and the reverse primers for FT were 5’-CTGGAACAACCTTTGGCAAT-3’ and 5’-TACACTGTTTGCCTGCCAAG-3’, respectively. IPP2 expression was used as a normalization control (Imaizumi et al., 2005). The following program was used for PCR: 95°C for 90 seconds, 45 cycles of 95°C for 10 seconds, 55°C for 20 seconds, and 72°C for 20 seconds. Each PCR was repeated at least three times.

Measurement

Protocol

RT-PCR ()

Experimental conditions

Sawa2007

Growth on MS plates 3% sucrose for 10.0 days (Sawa2007).

Light

Type Duration (days) Cycle (h) Start Duration Spectrum Source Intensity
diurnal light 10 24 0:00 16:00 white LED 90

Temperature

Type Duration (days) Cycle (h) Base (°C) Warm (°C) Warm start Warm duration
constant temperature 10 24 22 -- -- --

Sawa2007

Growth on MS 3% sucrose for 1.0 days (Sawa2007).

Light

Type Duration (days) Cycle (h) Start Duration Spectrum Source Intensity
diurnal light 1 24 0:00 16:00 white LED 90

Temperature

Type Duration (days) Cycle (h) Base (°C) Warm (°C) Warm start Warm duration
constant temperature 1 24 22 -- -- --

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